The Spangle mutation was first discovered in the aviary of Australian breeder S Casagrande in 1971. Although this mutation in other species is known as edged, it was named Spangle in budgies due to how similar the markings looked to spangle pigeons. Thanks to the quality breeders and dominant inheritance of the mutation, offspring were bred quickly and healthily.
SF Spangle budgies have reverse-colored markings on their wings, with the feather being clear-colored and the edge being black. This extends to their flight feathers and tail as well, the latter of which is almost fully clear-colored. While not present in every Spangle, their cheek markings are typically a mix of blue and white. No dark markings are present on the neck and head, much like an Opaline. Interestingly, these features make Spangle a type of pied mutation.
DF Spangle budgies are far different from the SF forms, being almost completely clear-colored with a darker “collar” around their faces (pictured third and fourth). Unlike Dark-eyed Clears and Lutinos, they have dark eyes with white irises.
SF Spangles with darker coloration on their wings are called melanistic spangles (pictured fifth). When combined with other mutations, these melanistic Spangles can form some truly unique colors.
